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a

The Great Paper Chase

Due: 11/18/04*

[* Due date: Those in by 11/18 will be guaranteed feedback by 12/9. Those in after 11/18 will be processed as time allows. A 10-point/day penalty will be assessed, including weekend days. None will be accepted after Thanksgiving.]

INTRODUCTION

To begin your search, you may want to check out the library's database section. You may want to use Elsevier's Science Direct, Wiley Interscience, and Dialog's Biosis. If you want to just jump in, try PubMed:

http://www.ncbi.nlm.nih.gov

In addition to the sites above, check out additional links on the Links page. [Let me know if you find useful sites which should be added.]

You can save your selected search results in a log, as you did for the earlier computer exercises. This is handy for making a useful printed short list to take to the library, or for use when in other databases, accessing on-line journals, or requesting interlibrary copies.

Some questions are general enough to allow you latitude in your searches; others are specific and allow fewer options in obtaining your information. All tools for doing literature searches are open to you. If you don't know how to approach a particular question, see me. THE HONOR SYSTEM DOES APPLY. The point of this exercise is, in part, to learn how to do a literature search. So- do your own search. Carpooling for library field-trips for journal access, or sharing information on database use or on finding on-line journals, is certainly acceptable.

You have several choices in getting the actual papers. [Remember the first computer exercise?] You can use our library and other libraries, such as UCD and UCSF. You can access some on-line journals for free, some on-line journals have limited access, and some charge a fee. PubMed is now providing direct links to journals on-line, and the free ones are marked. In addition, our library is part of some consortia which allow us access to specific journals. Particularly useful are Elsevier and Wiley Interscience. Other resources for free access are NewJour [linked on the library's Journal & Database page] and Highwire Press: www.highwire.org. You can request documents at the library from journals not available locally or free electronically. You can order papers on-line, but this is usually the most expensive solution, if you have a significant list of articles. When in a real, not virtual, library, be kind- don't make other people's searches a royal pain- there is plenty of space for all. Replace references on the central re-file shelves for journals when you are finished with a volume. That way others may locate them more easily.

Citations of your references are required, unless otherwise stated. (Be precise! You can count on us looking up some of these. Try copy/pasting from your log to avoid typos.) An example of a reference citation is as follows:

Thatcher, E.F. and Gershwin, L.J., 1989. Colostral transfer of bovine immunoglobulin E and dynamics of serum IgE in calves. Veterinary Immunology and Immunopathology 20:325-334.

Group your reference citations for each question with your answers to that question. They may precede or follow your answers as you choose, but they MUST BE PRECISE. A general bibliography without citation notes within your answers is not acceptable. Use either a number or the first author, and year if necessary, within your answer to refer to your references.

Papers are to be typed in a standard font style [please avoid script styles]; 12 pt is preferred, 10 pt is tolerated. Clearly indicate which questions are being answered and collate them in order. [E.g.: question 1 before question 3; a before b.] Check to be sure all subsections are answered and all parts of questions are answered, or points will be lost. Page limit: 12, including answers and citations. Please- no report covers or binders. [Saves you $$ & saves us time.] Just a single staple in the upper left-hand corner will do fine. [NO paperclips!]

Although most would assume that a statement on plagiarism is neither required nor needed, I unfortunately discovered otherwise. I direct you attention to the University's catalogue for campus policy on plagiarism. I expect answers in your own words, not that of the journals cited, or others not cited. I expect some thought and synthesis of information, not a regurgitation of phrases quoted. Quotes are permissible in appropriate circumstances and when referenced, but they should be used sparingly, if at all. Use your log appropriately. You can use it to collect papers and references. It is one useful way of organizing citation lists, which can then be copied to your paper chase file. But don't use it as a source of transferred blocks of text into your answers. If I find an answer is plagiarized, it will receive NO CREDIT. For help on this, check out the following:

http://www.indiana.edu/~wts/wts/plagiarism.html

If you find writing a problem, or you have difficulty organizing notes into coherent text, the Writing Center can be a lifesaver. They are very good at cutting to the chase of what the problem is and offering workable solutions in a minimum of time. [But don’t wait until the last minute.]

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THE QUESTIONS

PART A. Choose either question 1 or 2 below. [30 pt]

1. Intercellular collaborations resulting in cell signaling are a necessary part of any immune response. Cells need to interact with each other in specific ways, including direct contact through membrane-bound receptor-ligand interactions and/or soluble signal molecules, such as hormones and cytokines. Initial events in cell signaling lead to signal transduction, which as a term, includes events beginning with receipt of signal at the cell surface by whatever means, through intracellular signal pathways and second messengers, to affecting a response via gene activation and expression.

a. Integrins are an important group of adhesion molecules involved during cell migration and homing to specific tissues. Timothy Springer has been studying the cell signaling events associated with integrin binding and the specific molecular mechanisms involved. Describe briefly the methods used in his laboratory to investigate the functional and structural aspects of integrins during adhesion events. Summarize what has been discovered to date in terms of molecular conformational changes of integrins during adhesion and their role in cell signaling.

b. Jason Cyster is interested in chemokines and receptor-ligand interactions involved in lymphocyte homing in splenic white pulp, peripheral lymph nodes, and Peyer's patches. Describe what is understood about chemokine receptor signaling in B cell homing.

c. Phosphoinositide 3-kinases are involved in chemotaxis of both neutrophils and lymphocytes. Summarize what is understood to this point about the role and function of phosphoinositide 3-kinases.

2. Multiple pathways are involved in lymphocyte development in primary lymphoid tissues, including commitment toward B-cell or T-cell lineage and immunoglobulin or T-cell receptor rearrangement. During these processes, cells undergo both negative and positive selection prior to the release of virgin lymphocytes to the secondary lymphoid organs and tissues.

a. Ana Cumano investigates early lymphocyte development in fetal liver,  bone marrow, and thymus. She is involved in elucidating the different requirements and pathways found in these tissues relating to B cell and T cell maturation. Describe briefly the methods used to examine the immunoglobulin heavy chain rearrangements in cells located in the fetal live and bone marrow. Summarize the significance of her findings.

b. E2A gene products are involved in both B cell and thymocyte development. Cornelius Murre is interested in the regulation and function of these helix-loop-helix proteins. Describe what is known about the role of these E-proteins in development of both B cell and T cell lineages.

c. What is Notch-1? How is it involved in signaling during lymphocyte development and what is its significance?

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B. Choose either question 3 or 4 below. [40 pt]

3. There are many molecular mediators involved in host defense, both as innate responses to pathogenic challenges and in concert with adaptive immune responses. The following questions focus on only a few key areas of research.

a. Tumor necrosis factor [TNF] is a member of a large superfamily of proteins. Summarize the range of functions found for members of the TNF superfamily. Choose one of interest to you and describe the latest understanding of its mechanism of action and the specific role(s) ascribed to it.

b. The receptors for TNF ligands and co-receptors also make up another superfamily of proteins. CD40 is one such protein and appears to be involved in apoptosis and in the development of memory B cells rather than plasma cells. Describe what is known about CD40 and the mechanisms behind its involvement in cell death on one hand and generation of memory cells on the other.

c. Type I interferons, IFN-a and IFN-b, are well known as antiviral mediators. They are also involved in other types of activity, including innate response to non-viral pathogens.  Summarize the recent understandings of their role in non-viral responses and the mechanism of their functions.

d. Anti-microbial peptides are a diverse group of small molecules found widely in nature, including outside of the animal kingdom. They are significant in innate immunity. Cite a specific example and describe its mode of action and potential for use as a therapeutic agent in controlling infectious diseases.

4. The "War on Cancer" was declared during the Vietnam War, with statements proclaiming that a cure for cancer would be found within five years. The Vietnam War has been over for a long time, yet the fight against cancer rages on. The more we understand about immune responses and the mechanisms involved, the more we understand why the fight against cancer is so difficult.

a. T cell responses, particularly CTLs, are critical in the control of tumors. Many strategies have been explored to expand these responses. Novel vaccines have been developed against identified tumor antigens: however, these same antigens are often normal self-proteins being expressed on other types of cells. Cite a specific example of a tumor antigen that has been found to be a normal protein. What are the consequences when this antigen is used as a vaccine, both in terms of tumor control and in terms of side effects?

b. Although the focus has long been on MHC class I-restricted T cell responses, newer evidence is showing the importance of MHC class II-restricted antigen recognition. Briefly summarize the significance of processing exogenous antigen by CD4+ T cells and the relation to tumor responses. Discuss how this information could be used to develop improved treatments for tumors.

c. It has been found that low-avidity self-specific T cells survive the negative selection process in the thymus and normally do not proliferate. These cells have been linked to the development of autoimmune diseases; however, they are also attracting attention in terms of exploitation for anti-tumor therapies. Summarize the findings on these cells and their potential role in controlling tumors. After weighing the benefits and risks, what is your opinion about the efficacy of using this approach?

d. Antibodies are also produced in response to tumors. Describe a specific example of an immunoconjugate [antibody + toxin or other effecter molecule] and how it can used to augment treatments against some tumors. What is the current thinking about the utility of this type of approach?

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YOUR QUESTION

C. Design a question. [30 points] You write a question concerning an area of your interest. I review it, making comments or adding/subtracting parts or whatever. Then you do the search to answer your question.

I am frequently asked if the paper chosen for journal discussion can be connected to this part of the Paper Chase. In short- yes. [Can they be unrelated? Certainly.]

a. The question. Turn in by 10/14. I will return it by 10/19.

b. The answer. Due on 11/18 along with the rest of the answers.

NOTE: Be sure to include your original question, along with written comments,with your answers. [Don't assume I'll remember everything about your question out of the blue.]

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 Updated 10/4/04 by thatcher@sonoma.edu